Just got an email from the people at tweetjukebox.com
After finally getting some answers from our conversations with Twitter, and doing some research, we're confident we've found the cause of most of the 226 errors. There are large numbers of free accounts being opened and then used in an abusive fashion.
To combat this, effective immediately, we are going to institute a five tweet per day limit for FREE accounts. While I'm sure there are many folks who will take objection to this, our priority is that the platform remain in good standing with Twitter, and that paid accounts are in no way compromised by the actions of others.
It seems people will do anything if a dollar is dangled in front of them.